Invasion phenomena for heterogeneous reaction-diffusion equations are contemporary and challenging questions in applied mathematics. In this paper we are interested in the question of spreading for a reaction-diffusion equation when the subdomain where the reaction term is positive is shifting/contracting at a given speed $c$. This problem arises in particular in the modelling of the impact of climate change on population dynamics. By placing ourselves in the appropriate moving frame, this leads us to consider a reaction-diffusion-advection equation with a heterogeneous in space reaction term, in dimension $N\geq1$. We investigate the behaviour of the solution $u$ depending on the value of the advection constant~$c$, which typically stands for the velocity of climate change. We find that, when the initial datum is compactly supported, there exists precisely three ranges for $c$ leading to drastically different situations. In the lower speed range the solution always spreads, while in the upper range it always vanishes. More surprisingly, we find that that both spreading and vanishing may occur in an intermediate speed range. The threshold between those two outcomes is always sharp, both with respect to $c$ and to the initial condition. We also briefly consider the case of an exponentially decreasing initial condition, where we relate the decreasing rate of the initial condition with the range of values of~$c$ such that spreading occurs.
This is Part 2 of our work aimed at classifying the long-time behavior of the solution to a free boundary problem with monostable reaction term in space-time periodic media. In Part 1 (see \cite{ddl}) we have established a theory on the existence and uniqueness of solutions to this free boundary problem with continuous initial functions, as well as a spreading-vanishing dichotomy. We are now able to develop the methods of Weinberger \cite{w1, w2} and others \cite{fyz,lyz,lz1,lz2,lui} to prove the existence of asymptotic spreading speed when spreading happens, without knowing a priori the exist…
This paper is concerned with some spreading properties of monostable Lotka--Volterra two-species competition--diffusion systems when the initial values are null or exponentially decaying in a right half-line. Thanks to a careful construction of super-solutions and sub-solutions, we improve previously known results and settle open questions. In particular, we show that if the weaker competitor is also the faster one, then it is able to evade the stronger and slower competitor by invading first into unoccupied territories. The pair of speeds depends on the initial values. If these are null in a …
We establish in this article spreading properties for the solutions of equations of the type $\partial$ t u -- a(x)$\partial$ xx u -- q(x)$\partial$ x u = f (x, u), where a, q, f are only assumed to be uniformly continuous and bounded in x, the nonlinearity f is of monostable KPP type between two steady states 0 and 1 and the initial datum is compactly sup-ported. Using homogenization techniques, we construct two speeds w $\le$ w such that lim t$\rightarrow$+$\infty$ sup 0$\le$x$\le$wt |u(t, x)--1| = 0 for all w $\in$ (0, w) and lim t$\rightarrow$+$\infty$ sup x$\ge$wt |u(t, x)| = 0 for all w …
We consider a class of cooperative reaction-diffusion systems with free boundaries in one space dimension, where the diffusion terms are nonlocal, given by integral operators involving suitable kernel functions, and they are allowed not to appear in some of the equations in the system. The problem is monostable in nature, resembling the well known Fisher-KPP equation. Such a system covers various models arising from mathematical biology, with the Fisher-KPP equation as the simplest special case, where a "spreading-vanishing" dichotomy is known to govern the long time dynamical behaviour. The q…
We aim to classify the long-time behavior of the solution to a free boundary problem with monostable reaction term in space-time periodic media. Such a model may be used to describe the spreading of a new or invasive species, with the free boundary representing the expanding front. In time-periodic and space homogeneous environment, as well as in space-periodic and time autonomous environment, such a problem has been studied recently in \cite{dgp, dl}. In both cases, a spreading-vanishing dichotomy has been established, and when spreading happens, the asymptotic spreading speed is proved to ex…
We consider a multidimensional monostable reaction-diffusion equation whose nonlinearity involves periodic heterogeneity. This serves as a model of invasion for a population facing spatial heterogeneities. As a rescaling parameter tends to zero, we prove the convergence to a limit interface, whose motion is governed by the minimal speed (in each direction) of the underlying pulsating fronts. This dependance of the speed on the (moving) normal direction is in contrast with the homogeneous case and makes the analysis quite involved. Key ingredients are the recent improvement \cite{A-Gil} of the …
We focus on the spreading properties of solutions of monostable reaction-diffusion equations. Initial data are assumed to have heavy tails, which tends to accelerate the invasion phenomenon. On the other hand, the nonlinearity involves a weak Allee effect, which tends to slow down the process. We study the balance between the two effects. For algebraic tails, we prove the exact separation between "no acceleration and acceleration". This implies in particular that, for tails exponentially unbounded but lighter than algebraic , acceleration never occurs in presence of an Allee effect. This is in…
The theory of traveling waves and spreading speeds is developed for time-space periodic monotone semiflows with monostable structure. By using traveling waves of the associated Poincaré maps in a strong sense, we establish the existence of time-space periodic traveling waves and spreading speeds. We then apply these abstract results to a two species competition reaction-advection-diffusion model. It turns out that the minimal wave speed exists and coincides with the single spreading speed for such a system no matter whether the spreading speed is linearly determinate. We also obtain a set of s…
In this paper, we study the influence of an Allee effect on the spreading rate in a local reaction-diffusion-mutation equation modelling the invasion of cane toads in Australia. We are, in particular, concerned with the case when the diffusivity can take unbounded values. We show that the acceleration feature that arises in this model with a Fisher-KPP, or monostable, non-linearity still occurs when this non-linearity is instead bistable, despite the fact that this kills the small populations. This is in stark contrast to the work of Alfaro, Gui-Huan, and Mellet-Roquejoffre-Sire in related mod…
We study the asymptotic behaviour of solutions to the delayed monostable equation $(*)$: $u_{t}(t,x) = u_{xx}(t,x) - u(t,x) + g(u(t-h,x)),$ $x \in R,\ t >0,$ with monotone reaction term $g: R_+ \to R_+$. Our basic assumption is that this equation possesses pushed traveling fronts. First we prove that the pushed wavefronts are nonlinearly stable with asymptotic phase. Moreover, combinations of these waves attract, uniformly on $R$, every solution of equation $(*)$ with the initial datum sufficiently rapidly decaying at one (or at the both) infinities of the real line. These results provide a sh…
Many species see their range shifted poleward in response to global warming and need to keep pace in order to survive. To understand the effect of climate change on species ranges and its consequences on population dynamics, we consider a space-time heterogeneous reaction-diffusion equation in dimension 1, whose unknown~$u (t,x)$ stands for a population density. More precisely, the environment consists of two patches moving with a constant climate shift speed $c \geq 0$: in the invading patch $\{ t >0 , \, x \in \R \, | \ x < ct \}$ the growth rate is negative and, in the receding patch $\{ t …
This paper considers the spreading speed of cooperative nonlocal dispersal system with irreducible reaction functions and non-uniform initial data. Here the non-uniformity means that all components of initial data decay exponentially but their decay rates are different. It is well-known that in a monostable reaction-diffusion or nonlocal dispersal equation, different decay rates of initial data yield different spreading speeds. In this paper, we show that due to the cooperation and irreducibility of reaction functions, all components of the solution with non-uniform initial data will possess a…
In this paper, we extend and complement previous works about propagation in kinetic reaction-transport equations. The model we study describes particles moving according to a velocity-jump process, and proliferating according to a reaction term of monostable type. We focus on the case of bounded velocities, having dimension higher than one. We extend previous results obtained by the first author with Calvez and Nadin in dimension one. We study the large time/large scale hyperbolic limit via an Hamilton-Jacobi framework together with the half-relaxed limits method. We deduce spreading results a…
We devise a new geometric approach to study the propagation of disturbance - compactly supported data - in reaction diffusion equations. The method builds a bridge between the propagation of disturbance and of almost planar solutions. It applies to very general reaction-diffusion equations. The main consequences we derive in this paper are: a new proof of the classical Freidlin-Gartner formula for the asymptotic speed of spreading for periodic Fisher-KPP equations, extension of the formula to the monostable, combustion and bistable cases, existence of the asymptotic speed of spreading for equa…
We study nonlinear diffusion problems of the form $u_t=u_{xx}+f(u)$ with free boundaries. Such problems may be used to describe the spreading of a biological or chemical species, with the free boundary representing the expanding front. For special $f(u)$ of the Fisher-KPP type, the problem was investigated by Du and Lin [8]. Here we consider much more general nonlinear terms. For any $f(u)$ which is $C^1$ and satisfies $f(0)=0$, we show that the omega limit set $ω(u)$ of every bounded positive solution is determined by a stationary solution. For monostable, bistable and combustion types of non…
We focus on the spreading properties of solutions of monostable equations with fast diffusion. The nonlinear reaction term involves a weak Allee effect, which tends to slow down the propagation. We complete the picture of [3] by studying the subtle case where acceleration does occur and is induced by a combination of fast diffusion and of reactive growth. This requires the construction of new elaborate sub and supersolutions thanks to some underlying self-similar solutions.
We consider a multidimensional reaction-diffusion equation of either ignition or monostable type, involving periodic heterogeneity, and analyze the dependence of the propagation phenomena on the direction. We prove that the (minimal) speed of the underlying pulsating fronts depends continuously on the direction of propagation, and so does its associated profile provided it is unique up to time shifts. We also prove that the spreading properties \cite{Wein02} are actually uniform with respect to the direction.
In this paper, we study the existence and stability of travelling wave solutions of a kinetic reaction-transport equation. The model describes particles moving according to a velocity-jump process, and proliferating thanks to a reaction term of monostable type. The boundedness of the velocity set appears to be a necessary and sufficient condition for the existence of positive travelling waves. The minimal speed of propagation of waves is obtained from an explicit dispersion relation. We construct the waves using a technique of sub- and supersolutions and prove their \eb{weak} stability in a we…
We study the dispersion of a particle whose motion dynamics can be described by a forced velocity jump process. To investigate large deviations results, we study the Chapman-Kolmogorov equation of this process in the hyperbolic scaling (t,x,v) -> (t/epsilon,x/epsilon,v) and then, perform a Hopf-Cole transform which gives us a kinetic equation on a potential. We prove the convergence of this potential to the solution of a Hamilton-Jacobi equation. The hamiltonian can have a C1 singularity, as was previously observed in this kind of studies. This is a preliminary work before studying spreading r…
In this paper we consider a free boundary problem which models the spreading of an invasive species whose spreading is enhanced by the changing climate. We assume that the climate is shifting with speed c and obtain a complete classification of the long-time dynamical behaviour of the species. The model is similar to that in [9] with a slight refinement in the free boundary condition. While [9], like many works in the literature, investigates the case that unfavourable environment is shifting into the favourable habitat of the concerned species, here we examine the situation that the unfavoura…
Variation in genotypes may be responsible for differences in dispersal rates, directional biases, and growth rates of individuals. These traits may favor certain genotypes and enhance their spatio-temporal spreading into areas occupied by the less advantageous genotypes. We study how these factors influence the speed of spreading in the case of two competing genotypes and show that under the assumption of maintenance of spatially homogeneous total population the dynamics of the frequency of one of the genotypes is approximately described by the generalized Fisher-Kolmogorov-Petrovskii-Piskunov…
We focus on the spreading properties of solutions of monostable equations with non-linear diffusion. We consider both the porous medium diffusion and the fast diffusion regimes. Initial data may have heavy tails, which tends to accelerate the invasion phenomenon. On the other hand, the nonlinearity may involve a weak Allee effect, which tends to slow down the process. We study the balance between these three effects (nonlin-ear diffusion, initial tail, KPP nonlinearity/Allee effect), revealing the separation between "no acceleration" and "acceleration". In most of the cases where acceleration …
We investigate in this paper propagation phenomena for the heterogeneous reaction-diffusion equation $\partial_t u -Δu = f(t,u)$, $x\in R^N$, $t\in\R$, where f=f(t,u) is a KPP monostable nonlinearity which depends in a general way on t. A typical f which satisfies our hypotheses is f(t,u)=m(t) u(1-u), with m bounded and having positive infimum. We first prove the existence of generalized transition waves (recently defined by Berestycki and Hamel, Shen) for a given class of speeds. As an application of this result, we obtain the existence of random transition waves when f is a random stationary…
We consider the equation $u_t=u_{xx}+u_{yy}+b(x)f(u)+g(u)$, $(x,y)\in\mathbb R^2$ with monostable nonliearity, where $b(x)$ is a nonnegative measure on $\mathbb R$ that is periodic in $x.$ In the case where $b(x)$ is a smooth periodic function, there exists a pulsating travelling wave that propagates in the direction $(\cosθ,\sinθ)$ -- with average speed $c$ if and only if $c\geq c^*(θ,b),$ where $c^*(θ,b)$ is a certain positive number depending on $b.$ Moreover, the quantity $w(θ;{b})=\min_{|θ-ϕ|<\fracπ{2}}c^*(ϕ;{b})/\cos(θ-ϕ)$ is called the spreading speed. This theory can be extended by sho…
We study a nonlinear diffusion equation of the form $u_t=u_{xx}+f(u)\ (x\in [g(t),h(t)])$ with free boundary conditions $g'(t)=-u_x(t,g(t))+α$ and $h'(t)=-u_x(t,g(t))-α$ for some $α>0$. Such problems may be used to describe the spreading of a biological or chemical species, with the free boundaries representing the expanding fronts. When $α=0$, the problem was recently investigated by \cite{DuLin, DuLou}. In this paper we consider the case $α>0$. In this case shrinking (i.e. $h(t)-g(t)\to 0$) may happen, which is quite different from the case $α=0$. Moreover, we show that, under certain condit…
