Mutual interference and prey refuge are important drivers of predator-prey dynamics. The "exponent" or degree of mutual interference has been under much debate in theoretical ecology. In the present work, we investigate the interplay of the mutual interference exponent, on the behavior of a predator-prey model with a generalized Holling type functional response. We investigate stability properties of the system and derive conditions for the occurrence of saddle-node and Hopf-bifurcations. A sufficient condition for extinction of the prey species has also been derived for the model. In addition, we investigate the effect of a prey refuge on the population dynamics of the model and derive conditions for the prey refuge that would yield persistence of populations. We provide additional verification our analytical results via numerical simulations. Our findings are in accordance with classical experimental results in ecology by Gauss G. F (1934), that show that extinction of predator and prey populations is possible in a finite time period - but that bringing in refuge can effectively cause persistence.
A new model to investigate environmental effects of genetically distinguishable predators is presented. The Holling type II response function, modelling feeding satiation, leads to persistent system's oscillations, as in classical population models. An almost complete classification of the cases arising in the Routh-Hurwitz stability conditions mathematically characterizes the paper. It is instrumental as a guideline in the numerical experiments leading to the findings on the limit cycles. This result extends what found in an earlier parallel investigation containing a standard bilinear respon…
In this article, a Leslie-Gower Holling type III predator-prey model with disease in predator has been developed from both biological and mathematical point of view. The total population is divided into three classes, namely, prey, susceptible predator and infected predator. The local stability, global stability together with sufficient conditions for persistence of the ecosystem near biologically feasible equilibria is thoroughly investigated. Boundedness and existence of the system are established. All the important analytical findings are numerically verified using program software MATLAB a…
We discuss the stability and bifurcation analysis for a predator-prey system with non-linear Michaelis-Menten prey harvesting. The existence and stability of possible equilibria are investigated. We provide rigorous mathematical proofs for the existence of Hopf and saddle node bifurcations. We prove that the system exhibits Bogdanov-Takens bifurcation of codimension two, calculating the normal form. We provide several numerical simulations to illustrate our theoretical findings.
The goal of this paper is to explain how to derive the classical Rosenzweig-MacArthur's model by using a model with two groups of predators in which we can separate the vital dynamic and consumption of prey to describe the behavior of the predators. This will be especially very convenient if we want to add an age or size structure to the predator population. As mentioned by Holling (without mathematical model), we divide the population of predators into the searching and the handling predators. In this article we study some properties of this model and conclude the paper proving that the model…
We study a modified version of a prey-predator system with modified Leslie-Gower and Holling type II functional response studied by M.A. Aziz-Alaoui and M. Daher-Okiye. The modification consists in incorporating a refuge for preys, and substantially complicates the dynamics of the system. We study the local and global dynamics and the existence of cycles. We also investigate conditions for extinction or existence of a stationary distribution, in the case of a stochastic perturbation of the system.
In this manuscript, we consider temporal and spatio-temporal modified Holling-Tanner predator-prey models with predator-prey growth rate as a logistic type, Holling type II functional response and alternative food sources for the predator. From our result of the temporal model, we identify regions in parameter space in which Turing instability in the spatio-temporal model is expected and we show numerical evidence where the Turing instability leads to spatio-temporal periodic solutions. Subsequently, we analyse these instabilities. We use simulations to illustrate the behaviour of both the tem…
A predator prey system is investigated in this research, which is based on a modified version of the Leslie Gower scheme and a Holling-type II scheme with time dependent delays. Using Schauder's fixed point theorem, we studied the existence of pseudo almost periodic solution for the suggested model. Based on the suitable Lyapunov functional, sufficient conditions are established for the globally attractive pseudo almost periodic solution. At the end, two numerical examples are presented to demonstrate the effectiveness of our results.
This paper investigates a dynamical predator-prey interaction model that incorporates: (a) hunting cooperation among predators; (b) Allee effect in prey. We show all possible boundary and interior solutions. In order to analyze the stability of the solution, we make use of the Jacobian matrix and the resultant characteristic polynomial. Particularly, the sign of the eigenvalue is used to determine the stability of a solution. We then provide proof for stability of the interior solution. Finally, we verify our results numerically in MATLAB by plotting: (1) predator-prey intersection graphs; (2)…
In the present study, we have modified the traditional May-Holling-Tanner predator-prey model used to represent the interaction between least weasel and field-vole population by adding an Allee effect (strong and weak) on the field-vole population and alternative food source for the weasel population. It is shown that the dynamic is different from the original May-Holling-Tanner predator-prey interaction since new equilibrium points have appeared in the first quadrant. Moreover, the modified model allows the extinction of both species when the Allee effect (strong and weak) on the prey is incl…
Foraging movements of predator play an important role in population dynamics of prey-predator interactions, which have been considered as mechanisms that contribute to spatial self-organization of prey and predator. In nature, there are many examples of prey-predator interactions where prey is immobile while predator disperses between patches non-randomly through different factors such as stimuli following the encounter of a prey. In this work, we formulate a Rosenzweig-MacArthur prey-predator two patch model with mobility only in predator and the assumption that predators move towards patches…
The population dynamics of predator-prey systems in the presence of patch-specific predators are explored in a setting where the prey population has access to both habitats. The emphasis is in situations where patch-prey abundance drives prey-dispersal between patches, with the fragile prey populations, that is, populations subject to the Allee effect. The resulting four-dimensional model's mathematical analysis is carried out via sub-models that focus in lower dimensional settings. The outcomes depend on, and in fact they are quite sensitive to, the structure of the system, the range of param…
A delayed, discrete-time, prey-predator model with Allee effects imposed on prey and predator populations is defined, and dynamics of the system is characterized computationally. The parametric conditions for local asymptotic stability of equilibria are investigated with several illustrating examples. In addition, several periodic solutions were achieved. Furthermore, high degree of fractality of the prey-predator population trajectories is observed when the system is sufficiently delayed. This system has turned out to be a cooperative system, which is depicted using fractal dimension.
We consider a nonautonomous eco-epidemiological model with general functions for predation on infected and uninfected preys as well as general functions associated to the vital dynamics of the susceptible prey and predator populations. We obtain persistence and extinction results for the infected prey based on assumptions on systems related to the dynamics in the absence of infected preys. We apply our results to eco-epidemiological models constructed from predator-prey models existent in the literature. Some illustrative simulation is undertaken.
We analyse a modified Holling-Tanner predator-prey model where the predation functional response is of Holling type II and we incorporate a strong Allee effect associated with the prey species production. The analysis complements results of previous articles by Saez and Gonzalez-Olivares (SIAM J. Appl. Math. 59 1867-1878, 1999) and Arancibia-Ibarra and Gonzalez-Olivares (Proc. CMMSE 2015 130-141, 2015)discussing Holling-Tanner models which incorporate a weak Allee effect. The extended model exhibits rich dynamics and we prove the existence of separatrices in the phase plane separating basins o…
We analyse a modified May-Holling-Tanner predator-prey model considering an Allee effect in the prey and alternative food sources for predator. Additionally, the predation functional response or predation consumption rate is linear. The extended model exhibits rich dynamics and we prove the existence of separatrices in the phase plane separating basins of attraction related to oscillation, co-existence and extinction of the predator-prey population. We also show the existence of a homoclinic curve that degenerates to form a limit cycle and discuss numerous potential bifurcations such as saddle…
In this paper, we consider the dynamics of a predator-prey system of Gause type with cooperative hunting among predators and Holling III functional response. The known work numerically shows that the system exhibits saddle-node and Hopf bifurcations except homoclinic bifurcation for some special parameter values. Our results show that there are a weak focus of multiplicity three and a degenerate equilibrium with double zero eigenvalues (i.e., a cusp of codimension two) for general parameter conditions and the system can exhibit various bifurcations as perturbing the bifurcation parameters appr…
We study a predator-prey model with Holling type I functional response, an alternative food source for the predator, and multiple Allee effects on the prey. We show that the model has at most two equilibrium points in the first quadrant, one is always a saddle point while the other can be a repeller or an attractor. Moreover, there is always a stable equilibrium point that corresponds to the persistence of the predator population and the extinction of the prey population. Additionally, we show that when the parameters are varied the model displays a wide range of different bifurcations, such a…
We show that a number of models in virus dynamics, epidemiology and plant biology can be presented as ``damped" versions of the Lotka-Volterra predator-prey model, by analogy to the damped harmonic oscillator. The analogy deepens with the use of Lyapunov functions, which allow us to characterize their dynamics and even make some estimates.
A general framework for age-structured predator-prey systems is introduced. Individuals are distinguished into two classes, juveniles and adults, and several possible interactions are considered. The initial system of partial differential equations is reduced to a system of (neutral) delay differential equations with one or two delays. Thanks to this approach, physically correct models for predator-prey with delay are provided. Previous models are considered and analysed in view of the above results. A Rosenzweig-MacArthur model with delay is presented as an example.
A deterministic two-species predator-prey model with prey herd behavior is considered incorporating mutual interference and the effect of fear. We provide guidelines to the dynamical analysis of biologically feasible equilibrium points. We give conditions for the existence of some local and global bifurcations at the coexistence equilibrium. We also show that fear can induce extinction of the prey population from a coexistence zone in finite time. Our numerical simulations reveal that varying the strength of fear of predators with suitable choice of parameters can stabilize and destabilize the…
Dynamics and Control of Additional Food Provided Prey-Predator Systems exhibiting Holling Type-III Functional Response and Intra-specific Competition among Predators
The present paper deals with a prey-predator model with prey refuge proportion to both species and independent harvesting of each species. Our study shows that using refuge as control, it can break the limit circle of the system and reach the required state of equilibrium level. It is established the optimal harvesting policy. The boundedness, feasibility of interior equilibria, bionomic equilibrium have been determined. The main observation is that coefficient of refuge plays an important role in regulating the dynamics of the present system. Moreover the variation of the coefficient of refug…
We consider a predator-prey population model with prey gathering together for defense purposes. A transmissible unrecoverable disease affects the prey. We characterize the system behavior, establishing that ultimately either only the susceptible prey survive, or the disease becomes endemic, but the predators are wiped out. Another alternative is that the disease is eradicated, with sound prey and predators thriving at an equilibrium or through persistent population oscillations. Finally, the populations can thrive together, with the endemic disease. The only impossible alternative in these cir…
In this investigation we study the effects of climate change on predator-prey population dynamics. The interaction of predator and prey is described by the Variable Territory (VT) model with Allee effects in an Ordinary Differential Equation (ODE) framework. Climate influence is modeled in this ODE framework as an exogenous driver which affects the prey growth rate. Mathematically, this exogenous driver is a function of time, the climate function, that describes how favorable is the climate, on a scale from $-1$ to $+1$. Climate change has a number of effects; in this work, we focus on changes…
This paper deals with ratio-dependent predator-prey systems with delay. We will investigate under what conditions delay cannot cause instability in higher dimension. We give an example when delay causes instability.
Global Dynamics and Time-Optimal Control Studies for Additional Food provided Holling Type-III Mutually Interfering Prey-Predator Systems with Applications to Pest Management
本文研究提供额外食物的 Holling III 型相互干扰猎物-捕食者系统，通过平衡点、全局稳定性分析及以食物质量和数量为参数的时间最优控制，证明合适额外食物可使系统达到猎物消除状态。
We construct a global bifurcation diagram of the plane differential system $$ {l} \dot x = x(1-x)-x y/(a+x^2), \dot y = y(δ-βy/x), x(t)>0, y(t)>0, a>0, δ>0, β>0, $$ which describes the predator-prey interaction.
We prove the existence of infinitely many periodic solutions and complicated dynamics, due to the presence of a topological horseshoe, for the classical Volterra predator--prey model with a periodic harvesting. The proof relies on some recent results about chaotic planar maps combined with the study of geometric features which are typical of linked twist maps.
We consider a Rosenzweig-MacArthur predator-prey system which incorporates logistic growth of the prey in the absence of predators and a Holling type II functional response for interaction between predators and preys. We assume that parameters take values in a range which guarantees that all solutions tend to a unique limit cycle and prove estimates for the maximal and minimal predator and prey population densities of this cycle. Our estimates are simple functions of the model parameters and hold for cases when the cycle exhibits small predator and prey abundances and large amplitudes. The pro…
In this paper, we proposed a population model depicting the dynamics of a prey species showing group defence against a generalist predator. The group defence characteristic is represented by a non-monotonic functional response. We have established the local stability of the model around the co-existent equilibrium solution using a local Lyapunov function. Condition for existence Hopf bifurcation is obtained along with its normal form. Numerical simulations have been done to confirm the obtained analytical results as well as to validate the proposed model. Sensitivity analysis of the parameters…
We study the Bazykin predator-prey model with predator intraspecific interactions and ratio-dependent functional response and show the existence and stability of two interior equilibrium points. We prove that the model displays a wide range of different bifurcations, such as saddle-node bifurcations, Hopf bifurcations, homoclinic bifurcations and Bogdanov-Takens bifurcations. We use numerical simulations to further illustrate the impact changing the predator per capita consumption rate has on the basin of attraction of the stable equilibrium points, as well as the impact of changing the effici…
This paper is concerned with a predator-prey model of Leslie type with simplified Holling type IV functional response, provided that it has either a unique non-degenerate positive equilibrium or three distinct positive equilibria. The type and stability of each equilibrium, Hopf cyclicity of each weak focus, and the number and distribution of limit cycles in the first quadrant are studied. It is shown that every equilibrium cannot be a center. If system has a unique positive equilibrium which is a weak focus, then its order is at most $2$ and it has Hopf cyclicity $2$. Moreover, some sufficien…
A generalized seasonally-varying predator-prey model with Allee effect in the prey growth is investigated. The analysis is performed only on the basis of some properties determining the shape of the prey growth rate and the trophic interaction functions. General conditions for coexistence are determined, both in the case of weak and strong Allee effect. Finally, a modified Leslie-Gower predator-prey model with Allee effect is investigated. Numerical results illustrate the qualitative behaviors of the system, in particular the presence of periodic orbits.
We construct two ordinary-differential-equation models of a predator feeding adaptively on two prey types, and we evaluate the models' ability to fit data on freshwater plankton. We model the predator's switch from one prey to the other in two different ways: (1) smooth switching using a hyperbolic tangent function; and (2) by incorporating a parameter that changes abruptly across the switching boundary as a system variable that is coupled to the population dynamics. We conduct linear stability analyses, use approximate Bayesian computation (ABC) combined with a population Monte Carlo (PMC) me…
The dynamics of a prey-predator system with foraging facilitation among predators are investigated. The analysis involves the computation of many semi-algebraic systems of large degrees. We apply the pseudo-division reduction, real-root isolation technique and complete discrimination system of polynomial to obtain parameter conditions for the exact number of equilibria and their qualitative properties as well as a complete investigation of bifurcations including saddle-node, transcritical, pitchfork, Hopf and Bogdanov-Takens bifurcations. Moreover, numerical simulations are presented to suppor…
A predator-prey model with functional response Holling type II, Allee effect in the prey and a generalist predator is considered. It is shown that the model with strong Allee effect has at most two positive equilibrium point in the first quadrant, one is always a saddle point and the other exhibits multi-stability phenomenon since the equilibrium point can be stable or unstable. While the model with weak Allee effect has at most three positive equilibrium point in the first quadrant, one is always a saddle point and the other two can be stable or unstable node. In addition, when the parameters…
A general diffusive predator-prey model is investigated in this paper. We prove the global attractivity of constant equilibria when the conversion rate is small, and the non-existence of non-constant positive steady states when the conversion rate is large. The results are applied to several predator-prey models and give some ranges of parameters where complex pattern formation cannot occur.
A diffusive predator-prey system with predator interference and Neumann boundary conditions is considered in this paper. We derive some results on the existence and nonexistence of nonconstant stationary solutions. It is shown that there exist no nonconstant stationary solutions when the effect of the predator interference is strong or the conversion rate of the predator is large, and nonconstant stationary solutions emerge when the diffusion rate of the predator is large.
In this paper we present analytical solution of a fractional order predator-prey model, where prey grows logistically and predation occurs following type II response function, by homotopy perturbation method. Numerical solutions are presented to illustrate different particular cases. Our computational results show that accurate solution may be obtained with few iterations.
The diffusive Holling-Tanner predator-prey model with no-flux boundary conditions and nonlocal prey competition is considered in this paper. We show the existence of spatial nonhomogeneous periodic solutions, which is induced by nonlocal prey competition. In particular, the constant positive steady state can lose the stability through Hopf bifurcation when the given parameter passes through some critical values, and the bifurcating periodic solutions near such values can be spatially nonhomogeneous and orbitally asymptotically stable. This phenomenon is different from that in models without no…
Generalist predators consist an important component of an ecosystem which may act as a biocontrol agent and influence the dynamics significantly. In this paper, we have studied the effect of delayed logistic growth of the prey species with group defence behaviour. The Lyapunov stability criteria for the interior equilibrium point is derived. Also, the condition of Hopf-bifurcation and the point of bifurcation are obtained. The length of the delay is also estimated for the system to preserve stability. Numerical simulations are performed and illustrated to support the obtained analytical result…
A four dimensional ecoepidemiological model consisting of susceptible prey, infected prey, vaccinated prey and predator is formulated and analyzed in the present work. The functional response is assumed to be of Lotka-Volterra type. We studied systematically the behavior of the model with and without disease in prey. We analyzed mathematically the dynamics of the system such as boundedness of the solutions, existence and stability conditions of equilibria. The basic reproduction number \mathcal{R}_0 for the proposed model is computed. Disease is endemic if \mathcal{R}_0>1. Numerical simulation…
In this paper we consider a family of system with 2 predators feeding on one prey. We show how to construct a positively invariant set in which it is possible to define a Poincaré map for examining the behaviour of the system, mainly in the case when both predators survive. We relate it to examples from earlier works.
Following the well-extablished mathematical approach to persistence and its recent developments we give a rigorous theoretical explanation to the numerical results obtained for a certain prey-predator model with functional response of Holling type II equipped with an infectious disease in the predator population. The proof relies on some repelling conditions that can be applied in an iterative way on a suitable decomposition of the boundary. A full stability analysis is developed, showing how the "invasion condition" for the disease is derived. Some counterexamples and possible further investi…
We propose and study a two patch Rosenzweig-MacArthur prey-predator model with immobile prey and predator using two dispersal strategies. The first dispersal strategy is driven by the prey-predator interaction strength, and the second dispersal is prompted by the local population density of predators which is referred as the passive dispersal. The dispersal strategies using by predator are measured by the proportion of the predator population using the passive dispersal strategy which is a parameter ranging from 0 to 1. We focus on how the dispersal strategies and the related dispersal strengt…
Ratio-dependent predator-prey models have been increasingly favored by field ecologists where predator-prey interactions have to be taken into account the process of predation search. In this paper we study the conditions of the existence and stability properties of the equilibrium solutions in a reaction-diffusion model in which predator mortality is neither a constant nor an unbounded function, but it is increasing with the predator abundance. We show that analytically at a certain critical value a diffusion driven (Turing type) instability occurs, i.e. the stationary solution stays stable w…
In this note we present a study of the solutions associated to a particular spatial extension of the Rosenzweig-MacArthur model for predator and prey. The analysis presented here shows that positive steady state solutions emerge via a transcritical bifurcation mechanism, in accordance with the insight obtained from previous numerical and analytical results. In the model under discussion, prey is assumed to move avoiding crowds via a density-dependent diffusion and also incorporates the existence of a refuge zone, where predators cannot consume prey. Saturation in prey consumption is also inclu…
This paper deals with a free boundary problem of the Lotka-Volterra type prey-predator model with variable intrinsic growth rate for predator over a one dimensional habitat, in which the free boundary represents the spreading front and is caused only by the prey. In this problem it is assumed that the species can only invade further into the new environment from the right end of the initial region, and the spreading front expands at a speed that is proportional to the prey's population gradient at the front. Moreover, the variable intrinsic growth rate $r(t,x)$ is positive in pointwise but may…
A reaction-diffusion-advection predator-prey model with Holling type-II predator functional response is considered. We show the stability/instability of the positive steady state and the existence of a Hopf bifurcation when the diffusion and advection rates are large. Moreover, we show that advection rate can affect not only the occurrence of Hopf bifurcations but also the values of Hopf bifurcations.
We examine the interaction of transcritical and saddle-node bifurcations in a predator-prey-nutrient system that is stressed by the presence of a toxicant affecting the prey. This model, formulated by Kooi et al. ({\sl Ecol. Model.} {\bf 212}(2008), 304--318), has a two-dimensional invariant sub system with zero predator density. In the sub system, a pair of prey-nutrient equilibria is created in a saddle-node bifurcation, while predator invasion in modelled by a transcritical bifurcation of one of this pair. Interactions of these bifurcations at codimension-two points give rise to bistable, p…
