Modelling and analysis of a modified May-Holling-Tanner predator-prey model with Allee effect in the prey and an alternative food source for the predator
In the present study, we have modified the traditional May-Holling-Tanner predator-prey model used to represent the interaction between least weasel and field-vole population by adding an Allee effect (strong and weak) on the field-vole population and alternative food source for the weasel population. It is shown that the dynamic is different from the original May-Holling-Tanner predator-prey interaction since new equilibrium points have appeared in the first quadrant. Moreover, the modified model allows the extinction of both species when the Allee effect (strong and weak) on the prey is included, while the inclusion of the alternative food source for the predator shows that the system can support the coexistence of the populations, extinction of the prey and coexistence and oscillation of the populations at the same time. Furthermore, we use numerical simulations to illustrate the impact that changing the predation rate and the predator intrinsic growth rate have on the basin of attraction of the stable equilibrium point or stable limit cycle in the first quadrant. These simulations show the stabilisation of predator and prey populations and/or the oscillation of these two species over time.
A diffusive ratio-dependent Holling-Tanner system subject to Neumann boundary conditions is considered. The existence of multiple bifurcations, including Turing-Hopf bifurcation, Turing-Truing bifurcation, Hopf-double-Turing bifurcation and triple-Turing bifurcation, are given. Among them, the Turing-Hopf bifurcation are carried out in details by the normal form method. We theoretically prove that the system exists various spatio-temporal patterns, such as, non-constant steady state, the spatially inhomogeneous periodic or quasi-periodic solution, etc. Numerical simulations are presented to il…
We analyse a modified Holling-Tanner predator-prey model where the predation functional response is of Holling type II and we incorporate a strong Allee effect associated with the prey species production. The analysis complements results of previous articles by Saez and Gonzalez-Olivares (SIAM J. Appl. Math. 59 1867-1878, 1999) and Arancibia-Ibarra and Gonzalez-Olivares (Proc. CMMSE 2015 130-141, 2015)discussing Holling-Tanner models which incorporate a weak Allee effect. The extended model exhibits rich dynamics and we prove the existence of separatrices in the phase plane separating basins o…
The Turing-Hopf type spatiotemporal patterns in a diffusive Holling-Tanner model with discrete time delay is considered. A global Turing bifurcation theorem for $τ=0$ and a local Turing bifurcation theorem for $τ>0$ are given by the method of eigenvalue analysis and prior estimation. Further considering the degenerated situation, the existence of Bogdanov-Takens bifurcation and Turing-Hopf bifurcation are obtained. The normal form method is used to study the explicit dynamics near the Turing-Hopf singularity, and we establish the existence of various self-organized spatiotemporal patterns, suc…
We study a predator-prey model with Holling type I functional response, an alternative food source for the predator, and multiple Allee effects on the prey. We show that the model has at most two equilibrium points in the first quadrant, one is always a saddle point while the other can be a repeller or an attractor. Moreover, there is always a stable equilibrium point that corresponds to the persistence of the predator population and the extinction of the prey population. Additionally, we show that when the parameters are varied the model displays a wide range of different bifurcations, such a…
We analyse a modified May-Holling-Tanner predator-prey model considering an Allee effect in the prey and alternative food sources for predator. Additionally, the predation functional response or predation consumption rate is linear. The extended model exhibits rich dynamics and we prove the existence of separatrices in the phase plane separating basins of attraction related to oscillation, co-existence and extinction of the predator-prey population. We also show the existence of a homoclinic curve that degenerates to form a limit cycle and discuss numerous potential bifurcations such as saddle…
In this manuscript, we consider temporal and spatio-temporal modified Holling-Tanner predator-prey models with predator-prey growth rate as a logistic type, Holling type II functional response and alternative food sources for the predator. From our result of the temporal model, we identify regions in parameter space in which Turing instability in the spatio-temporal model is expected and we show numerical evidence where the Turing instability leads to spatio-temporal periodic solutions. Subsequently, we analyse these instabilities. We use simulations to illustrate the behaviour of both the tem…
In this paper, we discuss a hierarchical model, based on a Ricker competition model. The species considered are competing for resources and may be subject to an Allee effect due to mate limitation, anti-predator vigilance or aggression, cooperative predation or resource defense, or social thermoregulation. The species may be classified into a more dominant species and less dominant or ``wimpy" species or just as a predator and prey. The model under consideration also has components taking into account immigration in both species and a more structured Holling type II functional response. Local …
We prove the existence of traveling fronts in diffusive Rosenzweig-MacArthur and Holling-Tanner population models and investigate their relation with fronts in a scalar Fisher-KPP equation. More precisely, we prove the existence of fronts in a Rosenzweig-MacArthur predator-prey model in two situations: when the prey diffuses at the rate much smaller than that of the predator and when both the predator and the prey diffuse very slowly. Both situations are captured as singular perturbations of the associated limiting systems. In the first situation we demonstrate clear relations of the fronts wi…
The transition between strong and weak Allee effects in prey provides a simple regime shift in ecology. A deteriorating environment changes weak Allee effects into strong ones. In this paper, we study the interplay between the functional response of Holling type IV and both strong and weak Allee effects. The model investigated here presents complex dynamics and high codimension bifurcations. In particular, nilpotent cusp bifurcation, nilpotent saddle bifurcation and degenerate Hopf bifurcation of codimension 3 are completely analyzed, and the existence of homoclinic and heteroclinic loops are …
A predator-prey model with functional response Holling type II, Allee effect in the prey and a generalist predator is considered. It is shown that the model with strong Allee effect has at most two positive equilibrium point in the first quadrant, one is always a saddle point and the other exhibits multi-stability phenomenon since the equilibrium point can be stable or unstable. While the model with weak Allee effect has at most three positive equilibrium point in the first quadrant, one is always a saddle point and the other two can be stable or unstable node. In addition, when the parameters…
The diffusive Holling-Tanner predator-prey model with no-flux boundary conditions and nonlocal prey competition is considered in this paper. We show the existence of spatial nonhomogeneous periodic solutions, which is induced by nonlocal prey competition. In particular, the constant positive steady state can lose the stability through Hopf bifurcation when the given parameter passes through some critical values, and the bifurcating periodic solutions near such values can be spatially nonhomogeneous and orbitally asymptotically stable. This phenomenon is different from that in models without no…
In this paper, we consider a diffusive predator-prey system with spatial memory and predator-taxis. Since in this system, the memory delay appears in the diffusion term, and the diffusion term is nonlinear, the classical normal form of Hopf bifurcation in the reaction-diffusion system with delay can't be applied to this system. Thus, in this paper, we first derive an algorithm for calculating the normal form of Hopf bifurcation in this system. Then in order to illustrate the effectiveness of our newly developed algorithm, we consider the diffusive Holling-Tanner model with spatial memory and p…
