We consider a generalization of spatial branching coalescing processes in which the behaviour of individuals is not (necessarily) independent, on the contrary, individuals tend to take simultaneous actions. We show that these processes have moment duals, which happen to be multidimensional diffusions with jumps. Moment duality provides a general framework to study structural properties of the processes in this class. We present some conditions under which the expectation of the process is not affected by coordination and comment on the effect of coordination on the variance. We analyse several examples in more detail, including the nested coalescent, the peripatric coalescent with selection and coordinated migration, and the Parabolic Anderson Model.
We study several fundamental properties of a class of stochastic processes called spatial Lambda-coalescents. In these models, a number of particles perform independent random walks on some underlying graph G. In addition, particles on the same vertex merge randomly according to a given coalescing mechanism. A remarkable property of mean-field coalescent processes is that they may come down from infinity, meaning that, starting with an infinite number of particles, only a finite number remains after any positive amount of time, almost surely. We show here however that, in the spatial setting, …
We design a theoretic tree-based functional representation of a class of Feynman-Kac particle distributions, including an extension of the Wick product formula to interacting particle systems. These weak expansions rely on an original combinatorial, and permutation group analysis of a special class of forests. They provide refined non asymptotic propagation of chaos type properties, as well as sharp $\LL\_p$-mean error bounds, and laws of large numbers for $U$-statistics. Applications to particle interpretations of the top eigenvalues, and the ground states of Schrödinger semigroups are also d…
We present a coalescent process where three particles merge at each coagulation step. Using a random walk representation, we prove duality with a fragmentation process, whose fragmentation law we specify explicitly. Furthermore, we give a second construction of the coalescence in terms of random binary forests and study asymptotic properties. Starting from N particles of unit mass, we obtain under an appropriate rescaling when N tends to infinity a well-known binary coalescence, the so-called standard additive coalescent.
We examine the total number of collisions $C_n$ in the $Λ$-coalescent process which starts with $n$ particles. A linear growth and a stable limit law for $C_n$ are shown under the assumption of a power-like behaviour of the measure $Λ$ near 0 with exponent $0<α<1$.
Coalescent theory is the study of random processes where particles may join each other to form clusters as time evolves. These notes provide an introduction to some aspects of the mathematics of coalescent processes and their applications to theoretical population genetics and other fields such as spin glass models. The emphasis is on recent work concerning in particular the connection of these processes to continuum random trees and spatial models such as coalescing random walks.
Take a continuous-time Galton-Watson tree. If the system survives until a large time $T$, then choose $k$ particles uniformly from those alive. What does the ancestral tree drawn out by these $k$ particles look like? Some special cases are known but we give a more complete answer. We concentrate on near-critical cases where the mean number of offspring is $1+μ/T$ for some $μ\in\mathbb{R}$, and show that a scaling limit exists as $T\to\infty$. Viewed backwards in time, the resulting coalescent process is topologically equivalent to Kingman's coalescent, but the times of coalescence have an inte…
Consider a population evolving as a critical continuous-time Galton-Watson (GW) tree. Conditional on the population surviving until a large time $T$, sample $k$ individuals uniformly at random (without replacement) from amongst those alive at time $T$. What is the genealogy of this sample of individuals? In cases where the offspring distribution has finite variance, the probabilistic properties of the joint ancestry of these $k$ particles are well understood, as seen in \cite{HJR20, J19}. In the present article, we study the joint ancestry of a sample of $k$ particles under the following regim…
We introduce and analyze a novel type of coalescent processes called cross-multiplicative coalescent that models a system with two types of particles, $A$ and $B$. The bonds are formed only between the pairs of particles of opposite types with the same rate for each bond, producing connected components made of particles of both types. We analyze and solve the Smoluchowski coagulation system of equations obtained as a hydrodynamic limit of the corresponding Marcus-Lushnikov process. We establish that the cross-multiplicative kernel is a gelling kernel, and find the gelation time. As an applicat…
Let $\mathbb{T}^d_N$, $d\ge 2$, be the discrete $d$-dimensional torus with $N^d$ points. Place a particle at each site of $\mathbb{T}^d_N$ and let them evolve as independent, nearest-neighbor, symmetric, continuous-time random walks. Each time two particles meet, they coalesce into one. Denote by $C_N$ the first time the set of particles is reduced to a singleton. Cox [6] proved the existence of a time-scale $θ_N$ for which $C_N/θ_N$ converges to the sum of independent exponential random variables. Denote by $Z^N_t$ the total number of particles at time $t$. We prove that the sequence of Marko…
Let $Λ$ be a finite measure on the unit interval. A $Λ$-Fleming-Viot process is a probability measure valued Markov process which is dual to a coalescent with multiple collisions ($Λ$-coalescent) in analogy to the duality known for the classical Fleming Viot process and Kingman's coalescent, where $Λ$ is the Dirac measure in 0. We explicitly construct a dual process of the coalescent with simultaneous multiple collisions ($Ξ$-coalescent) with mutation, the $Ξ$-Fleming-Viot process with mutation, and provide a representation based on the empirical measure of an exchangeable particle system alon…
Consider a haploid population which has evolved through an exchangeable reproduction dynamics, and in which all individuals alive at time $t$ have a most recent common ancestor (MRCA) who lived at time $A_t$, say. As time goes on, not only the population but also its genealogy evolves: some families will get lost from the population and eventually a new MRCA will be established. For a time-stationary situation and in the limit of infinite population size $N$ with time measured in $N$ generations, i.e. in the scaling of population genetics which leads to Fisher-Wright diffusions and Kingman's c…
We describe a new general connection between $Λ$-coalescents and genealogies of continuous-state branching processes. This connection is based on the construction of an explicit coupling using a particle representation inspired by the lookdown process of Donnelly and Kurtz. This coupling has the property that the coalescent comes down from infinity if and only if the branching process becomes extinct, thereby answering a question of Bertoin and Le Gall. The coupling also offers new perspective on the speed of coming down from infinity and allows us to relate power-law behavior for $N^Λ(t)$ to …
This article introduces new tools to study self-organisation in a family of simple cellular automata which contain some particle-like objects with good collision properties (coalescence) in their time evolution. We draw an initial configuration at random according to some initial $σ$-ergodic measure $μ$, and use the limit measure to descrbe the asymptotic behaviour of the automata. We first take a qualitative approach, i.e. we obtain information on the limit measure(s). We prove that only particles moving in one particular direction can persist asymptotically. This provides some previously unk…
The nested Kingman coalescent describes the dynamics of particles (called genes) contained in larger components (called species), where pairs of species coalesce at constant rate and pairs of genes coalesce at constant rate provided they lie within the same species. We prove that starting from $rn$ species, the empirical distribution of species masses (numbers of genes$/n$) at time $t/n$ converges as $n\to\infty$ to a solution of the deterministic coagulation-transport equation $$ \partial_t d \ = \ \partial_x ( ψd ) \ + \ a(t)\left(d\star d - d \right), $$ where $ψ(x) = cx^2$, $\star$ denotes…
In this paper we show the existence of the minimal solution to the multidimensional Lambert-Euler inversion, a multidimensional generalization of $[-e^{-1} ,0)$ branch of Lambert W function $W_0(x)$. Specifically, for a given nonnegative irreducible symmetric matrix $V \in \mathbb{R}^{k \times k}$, we show that for ${\bf u}\in(0,\infty)^k$, if equation $$y_j \exp\{-{\bf e}_j^T V {\bf y} \} = u_j ~~~~~~\forall j=1,...,k,$$ has at least one solution, it must have a minimal solution ${\bf y}^*$, where the minimum is achieved in all coordinates $y_j$ simultaneously. Moreover, such ${\bf y}^*$ is t…
We revisit the discrete additive and multiplicative coalescents, starting with $n$ particles with unit mass. These cases are known to be related to some "combinatorial coalescent processes": a time reversal of a fragmentation of Cayley trees or a parking scheme in the additive case, and the random graph process $(G(n,p))_p$ in the multiplicative case. Time being fixed, encoding these combinatorial objects in real-valued processes indexed by the line is the key to describing the asymptotic behaviour of the masses as $n\to +\infty$. We propose to use the Prim order on the vertices instead of the…
The generalized Fleming-Viot processes were defined in 1999 by Donnelly and Kurtz using a particle model and by Bertoin and Le Gall in 2003 using stochastic flows of bridges. In both methods, the key argument used to characterize these processes is the duality between these processes and exchangeable coalescents. A larger class of coalescent processes, called distinguished coalescents, was set up recently to incorporate an immigration phenomenon in the underlying population. The purpose of this article is to define and characterize a class of probability-measure valued processes called the gen…
We consider a system of particles which perform branching Brownian motion with negative drift and are killed upon reaching zero, in the near-critical regime where the total population stays roughly constant with approximately N particles. We show that the characteristic time scale for the evolution of this population is of order $(\log N)^3$, in the sense that when time is measured in these units, the scaled number of particles converges to a variant of Neveu's continuous-state branching process. Furthermore, the genealogy of the particles is then governed by a coalescent process known as the …
We study the genealogy of a solvable population model with $N$ particles on the real line which evolves according to a discrete-time branching process with selection. At each time step, every particle gives birth to children around $a$ times its current position, where $a>0$ is a parameter of the model. Then, the $N$ rightmost new-born children are selected to form the next generation. We show that the genealogical trees of the process converge to those of a Beta coalescent as $N \to \infty$. The process we consider can be seen as a toy-model version of a continuous-time branching process with…
We present simple conditions under which the limiting genealogical process associated with a class of interacting particle systems with non-neutral selection mechanisms, as the number of particles grows, is a time-rescaled Kingman coalescent. Sequential Monte Carlo algorithms are popular methods for approximating integrals in problems such as non-linear filtering and smoothing which employ this type of particle system. Their performance depends strongly on the properties of the induced genealogical process. We verify the conditions of our main result for standard sequential Monte Carlo algorit…
The ancestral selection graph in population genetics was introduced by KroneNeuhauser (1997) as an analogue of the coalescent genealogy of a sample of genes from a neutrally evolving population. The number of particles in this graph, followed backwards in time, is a birth and death process with quadratic death and linear birth rates. In this paper an explicit form of the probability distribution of the number of particles is obtained by using the density of the allele frequency in the corresponding diffusion model obtained by Kimura (1955). It is shown that the process of fixation of the allel…
The goal of this paper is to prove rigorous results for the behavior of genealogies in a one-dimensional long range biased voter model introduced by Hallatschek and Nelson [25]. The first step, which is easily accomplished using results of Mueller and Tribe [38], is to show that when space and time are rescaled correctly, our biased voter model converges to a Wright-Fisher SPDE. A simple extension of a result of Durrett and Restrepo [18] then shows that the dual branching coalescing random walk converges to a branching Brownian motion in which particles coalesce after an exponentially distribu…
