Cooperation is a persistent behavioral pattern of entities pooling and sharing resources. Its ubiquity in nature poses a conundrum. Whenever two entities cooperate, one must willingly relinquish something of value to the other. Why is this apparent altruism favored in evolution? Classical solutions assume a net fitness gain in a cooperative transaction which, through reciprocity or relatedness, finds its way back from recipient to donor. We seek the source of this fitness gain. Our analysis rests on the insight that evolutionary processes are typically multiplicative and noisy. Fluctuations have a net negative effect on the long-time growth rate of resources but no effect on the growth rate of their expectation value. This is an example of non-ergodicity. By reducing the amplitude of fluctuations, pooling and sharing increases the long-time growth rate for cooperating entities, meaning that cooperators outgrow similar non-cooperators. We identify this increase in growth rate as the net fitness gain, consistent with the concept of geometric mean fitness in the biological literature. This constitutes a fundamental mechanism for the evolution of cooperation. Its minimal assumptions make it a candidate explanation of cooperation in settings too simple for other fitness gains, such as emergent function and specialization, to be probable. One such example is the transition from single cells to early multicellular life.
Game theory research on the snowdrift game has showed that gradual evolution of the continuously varying level of cooperation in joint enterprises can demonstrate evolutionary merging as well as evolutionary branching. However, little is known about the consequences of changes in diversity at the cooperation level. In the present study I consider effects of costly rewards on the continuous snowdrift game. I show that not evolutionary merging but evolutionary branching can promote the emergence of pool reward, which can then enhance social welfare.
We study the combined influence of selection and random fluctuations on the evolutionary dynamics of two-strategy ("cooperation" and "defection") games in populations comprising cooperation facilitators. The latter are individuals that support cooperation by enhancing the reproductive potential of cooperators relative to the fitness of defectors. By computing the fixation probability of a single cooperator in finite and well-mixed populations that include a fixed number of facilitators, and by using mean field analysis, we determine when selection promotes cooperation in the important classes …
Individuals might abstain from participating in an instance of an evolutionary game for various reasons, ranging from lack of interest to risk aversion. In order to understand the consequences of such diverse activity patterns on the evolution of cooperation, we study a weak prisoner's dilemma where each player's participation is probabilistic rather than certain. Players that do not participate get a null payoff and are unable to replicate. We show that inactivity introduces cascading failures of cooperation, which are particularly severe on scale-free networks with frequently inactive hubs. …
A binary game is introduced and analysed. N players have to choose one of the two sides independently and those on the minority side win. Players uses a finite set of ad hoc strategies to make their decision, based on the past record. The analysing power is limited and can adapt when necessary. Interesting cooperation and competition pattern of the society seem to arise and to be responsive to the payoff function.
Game theory deals with strategic interactions among players and evolutionary game dynamics tracks the fate of the players' populations under selection. In this paper, we consider the replicator equation for two-player-two-strategy games involving cooperators and defectors. We modify the equation to include the effect of mutation and also delay that corresponds either to the delayed information about the population state or in realizing the effect of interaction among players. By focusing on the four exhaustive classes of symmetrical games -- the Stag Hunt game, the Snowdrift game, the Prisoner…
Evolutionary mechanisms that promote cooperation may not promote social welfare
Teesside University(提赛德大学);University of Birmingham(伯明翰大学);University of Maribor(马里博尔大学);Community Healthcare Center Dr. Adolf Drolc Maribor(阿道夫·德罗尔奇博士马里博尔社区医疗中心);Complexity Science Hub Vienna(维也纳复杂科学中心);Kyung Hee University(庆熙大学)
Diffusion-induced instabilities promote cooperation in eco-evolutionary networks
Bagchi School of Public Health, Ahmedabad University(艾哈迈达巴德大学巴吉公共卫生学院);Department of Physical Sciences, Indian Institute of Science Education and Research Kolkata(印度科学教育与研究学院加尔各答分校物理科学系);Department of Mathematics and Namur Institute for Complex Systems - naXys, University of Namur(南纳尔大学数学与南纳尔复杂系统研究所-naXys);Faculty of Natural Sciences and Mathematics, University of Maribor(马里博尔大学自然科学与数学学院);Community Healthcare Center Dr. Adolf Drolc Maribor(马里博尔阿道夫·德罗尔克博士社区健康中心);Department of Physics, Kyung Hee University(庆熙大学物理系);University College, Korea University(韩国大学综合研究生院);Physics and Applied Mathematics Unit, Indian Statistical Institute(印度统计物理学与应用数学单元)
We compare the likelihood of different socially relevant features to allow the evolutionary emergence and maintenance of cooperation in a generalized variant of the iterated Prisoners Dilemma game. Results show that the average costs/benefit balance of cooperation is the primary constraint for its establishment and maintenance. Behavior increasing inclusive fitness such as assortation, homophily, kin-selection and tagging of individuals, is second in importance. Networks characteristics were the least important in favoring the establishment and maintenance of cooperation, despite being the mos…
It is well known that cooperation cannot be an evolutionary stable strategy for a non-iterative game in a well-mixed population. In contrast, structured populations favor cooperation since cooperators can benefit each other by forming local clusters. Previous studies have shown that scale-free networks strongly promote cooperation. However, little is known about the invasion mechanism of cooperation in scale-free networks. To study microscopic and macroscopic behaviors of cooperators' invasion, we conducted computational experiments of the evolution of cooperation in scale-free networks where,…
Large-scale cooperation underpins the evolution of ecosystems and the human society, and the collective behaviors by self-organization of multi-agent systems are the key for understanding. As artificial intelligence (AI) prevails in almost all branches of science, it would be of great interest to see what new insights of collective behavior could be obtained from a multi-agent AI system. Here, we introduce a typical reinforcement learning (RL) algorithm -- Q learning into evolutionary game dynamics, where agents pursue optimal action on the basis of the introspectiveness rather than the birth-…
Pro-social punishment is a key driver of harmonious and stable society. However, this institution is vulnerable to corruption since law-violators can avoid sanctioning by paying bribes to corrupt law-enforcers. Consequently, to understand how altruistic behavior survives in a corrupt environment is an open question. To reveal potential explanations here we introduce corrupt enforcers and violators into the public goods game with pool punishment, and assume that punishers, as corrupt enforcers, may select defectors probabilistically to take a bribe from, and meanwhile defectors, as corrupt viol…
In the framework of the evolutionary dynamics of the Prisoner's Dilemma game on complex networks, we investigate the possibility that the average level of cooperation shows hysteresis under quasi-static variations of a model parameter (the "temptation to defect"). Under the "discrete replicator" strategy updating rule, for both Erdos-Renyi and Barabasi-Albert graphs we observe cooperation hysteresis cycles provided one reaches tipping point values of the parameter; otherwise, perfect reversibility is obtained. The selective fixation of cooperation at certain nodes and its organization in coope…
The prisoner's dilemma has long been considered the paradigm for studying the emergence of cooperation among selfish individuals. Because of its importance, it has been studied through computer experiments as well as in the laboratory and by analytical means. However, there are important differences between the way a system composed of many interacting elements is simulated by a digital machine and the manner in which it behaves when studied in real experiments. In some instances, these disparities can be marked enough so as to cast doubt on the implications of cellular automata type simulatio…
We study the evolutionary Prisoner's Dilemma on two social networks obtained from actual relational data. We find very different cooperation levels on each of them that can not be easily understood in terms of global statistical properties of both networks. We claim that the result can be understood at the mesoscopic scale, by studying the community structure of the networks. We explain the dependence of the cooperation level on the temptation parameter in terms of the internal structure of the communities and their interconnections. We then test our results on community-structured, specifical…
We propose a model of emergence of cooperation in evolutionary games that high- lights the role of network formation and effect of network structure. In line with empirical data, the model proposes a mechanism that explains the persistence of heterogeneous types (heterogeneity in rules for changing behavior) within a population, and in particular the sustainability of altruism (presence of unconditional cooperators) even in case of strong social dilemma. This explanation constitutes an alternative to the choice and refusal mechanism that is often presented as an explanation for cooperation on …