The observed cooperation on the level of genes, cells, tissues, and individuals has been the object of intense study by evolutionary biologists, mainly because cooperation often flourishes in biological systems in apparent contradiction to the selfish goal of survival inherent in Darwinian evolution. In order to resolve this paradox, evolutionary game theory has focused on the Prisoner's Dilemma (PD), which incorporates the essence of this conflict. Here, we encode strategies for the iterated Prisoner's Dilemma (IPD) in terms of conditional probabilities that represent the response of decision pathways given previous plays. We find that if these stochastic strategies are encoded as genes that undergo Darwinian evolution, the environmental conditions that the strategies are adapting to determine the fixed point of the evolutionary trajectory, which could be either cooperation or defection. A transition between cooperative and defective attractors occurs as a function of different parameters such a mutation rate, replacement rate, and memory, all of which affect a player's ability to predict an opponent's behavior.
Stability of evolutionary dynamics of non-repeated Prisoner's Dilemma game with non-uniform interaction rates [1], via benefit and cost dilemma is studied . Moreover, the stability condition (b+c/b-c)2 < r1r3 is derived in case of coexistence between cooperators and defectors .If r1,r3 -> infinity cooperation is the dominant strategy and defectors can no longer exploit cooperators.
We investigate the evolution of cooperation on a non - growth network model with death/birth dynamics. Nodes reproduce under selection for higher payoffs in a prisoners dilemma game played between network neighbours. The mean field characteristics of the model are explored and an attempt is made to understand the size dependent behaviour of the model in terms of fluctuations in the strategy densities. We also briefly comment on the role of strategy mutation in regulating the strategy densties.
We investigate the evolutionary dynamics of an age-structured population under weak frequency-dependent selection. It turns out that the weak selection is affected in a non-trivial way by the life-history trait. We can disentangle the dynamics, based on the appearance of different time scales. These time scales, which seem to form a universal structure in the interplay of weak selection and life-history traits, allow us to reduce the infinite dimensional model to a one-dimensional modified replicator equation. The modified replicator equation is then used to investigate cooperation (the prison…
We seek a route to the equilibrium where all the agents cooperate in the iterated prisoner's dilemma game on a two-dimensional plane, focusing on the role of tit-for-tat strategy. When a time horizon, within which a strategy can recall the past, is one time step, an equilibrium can be achieved as cooperating strategies dominate the whole population via proliferation of tit-for-tat. Extending the time horizon, we filter out poor strategies by simplified replicator dynamics and observe a similar evolutionary pattern to reach the cooperating equilibrium. In particular, the rise of a modified tit-…
Evolution of cooperation in the prisoner's dilemma game is studied where initially all players are linked via a regular graph, having four neighbors each. Simultaneously with the strategy evolution, players are allowed to make new connections and thus permanently extend their neighborhoods, provided they have been successful in passing their strategy to the opponents. We show that this simple coevolutionary rule shifts the survival barrier of cooperators towards high temptations to defect and results in highly heterogeneous interaction networks with an exponential fit best characterizing their…
The world in which we are living is a huge network of networks and should be described by interdependent networks. The interdependence between networks significantly affects the evolutionary dynamics of cooperation on them. Meanwhile, due to the diversity and complexity of social and biological systems, players on different networks may not interact with each other by the same way, which should be described by multiple models in evolutionary game theory, such as the Prisoner's Dilemma and Snowdrift Game. We therefore study the evolutionary dynamics of cooperation on two interdependent networks…
Evolutionary game theory assumes that players replicate a highly scored player's strategy through genetic inheritance. However, when learning occurs culturally, it is often difficult to recognize someone's strategy just by observing the behaviour. In this work, we consider players with memory-one stochastic strategies in the iterated prisoner's dilemma, with an assumption that they cannot directly access each other's strategy but only observe the actual moves for a certain number of rounds. Based on the observation, the observer has to infer the resident strategy in a Bayesian way and chooses …
We examine the classical contents of quantum games. It is shown that a quantum strategy can be interpreted as a classical strategies with effective density-dependent game matrices composed of transposed matrix elements. In particular, successful quantum strategies in dilemma games are interpreted in terms of a symmetrized game matrix that corresponds to an altruistic game plan.
Dense cooperative networks are an essential element of social capital for a prosperous society. These networks enable individuals to overcome collective action dilemmas by enhancing trust. In many biological and social settings, network structures evolve endogenously as agents exit relationships and build new ones. However, the process by which evolutionary dynamics lead to self-organization of dense cooperative networks has not been explored. Our large group prisoner's dilemma experiments with exit and partner choice options show that core-periphery segregation of cooperators and defectors dr…
The Prisoner's Dilemma, a 2-person game in which the players can either cooperate or defect, is a common paradigm for studying the evolution of cooperation, when individuals exhibit variable degrees of cooperation. It is known that in the presence of spatial structure, when individuals ``play against'' their neighbours, and ``compare to'' them, cooperative investments can evolve to considerable levels. Here we examine the effect of increasing the neighbourhood size: we find that the mean-field limit of no cooperation is reached for a critical neighbourhood size of about five neighbours. We als…
Humans and other animals can adapt their social behavior in response to environmental cues including the feedback obtained through experience. Nevertheless, the effects of the experience-based learning of players in evolution and maintenance of cooperation in social dilemma games remain relatively unclear. Some previous literature showed that mutual cooperation of learning players is difficult or requires a sophisticated learning model. In the context of the iterated Prisoner's Dilemma, we numerically examine the performance of a reinforcement learning model. Our model modifies those of Karand…
In the framework of the paradigmatic prisoner's dilemma, we investigate the evolutionary dynamics of social dilemmas in the presence of "cooperation facilitators". In our model, cooperators and defectors interact as in the classical prisoner's dilemma game, where selection favors defection. However, here the presence of a small number of cooperation facilitators enhances the fitness (reproductive potential) of cooperators, while it does not alter that of defectors. In a finite population of size N, the dynamics of the prisoner's dilemma with facilitators is characterized by the probability tha…
Differences between men and women have intrigued generations of social scientists, who have found that the two sexes behave differently in settings requiring competition, risk taking, altruism, honesty, as well as many others. Yet, little is known about whether there are gender differences in cooperative behavior. Previous evidence is mixed and inconclusive. Here I shed light on this topic by analyzing the totality of studies that my research group has conducted since 2013. This is a dataset of 10,951 observations coming from 7,322 men and women living in the US, recruited through Amazon Mecha…
A non-local rare mutations model for quasispecies and Prisoner's dilemma: numerical assessment of qualitative behaviour
Università di Napoli “Parthenope”（那不勒斯帕尔特诺佩大学）；Istituto per le Applicazioni del Calcolo “M. Picone”, Consiglio Nazionale delle Ricerche（意大利国家研究委员会计算应用研究所）
Recent studies on the evolutionary dynamics of the Prisoner's Dilemma game in scale-free networks have demonstrated that the heterogeneity of the network interconnections enhances the evolutionary success of cooperation. In this paper we address the issue of how the characterization of the asymptotic states of the evolutionary dynamics depends on the initial concentration of cooperators. We find that the measure and the connectedness properties of the set of nodes where cooperation reaches fixation is largely independent of initial conditions, in contrast with the behavior of both the set of n…
Evolutionary Prisoner's Dilemma games with quenched inhomogeneities in the spatial dynamical rules are considered. The players following one of the two pure strategies (cooperation or defection) are distributed on a two-dimensional lattice. The rate of strategy adoption from a randomly chosen neighbors are controlled by the payoff difference and a two-value pre-factor $w$ characterizing the players whom the strategy learned from. The reduced teaching activity of players is distributed randomly with concentrations $ν$ at the beginning and fixed further on. Numerical and analytical calculations …
Recent experimental evidence [Grujić et al., PLoS ONE 5, e13749 (2010)] on the spatial Prisoner's Dilemma suggests that players choosing to cooperate or not on the basis of their previous action and the actions of their neighbors coexist with steady defectors and cooperators. We here study the coexistence of these three strategies in the multiplayer iterated Prisoner's Dilemma by means of the replicator dynamics. We consider groups with n = 2, 3, 4 and 5 players and compute the payoffs to every type of player as the limit of a Markov chain where the transition probabilities between actions are…
Tumor development is an evolutionary process in which a heterogeneous population of cells with differential growth capabilities compete for resources in order to gain a proliferative advantage. What are the minimal ingredients needed to recreate some of the emergent features of such a developing complex ecosystem? What is a tumor doing before we can detect it? We outline a mathematical model, driven by a stochastic Moran process, in which cancer cells and healthy cells compete for dominance in the population. Each are assigned payoffs according to a Prisoner's Dilemma evolutionary game where t…
We study the evolution of cooperation in the spatial prisoner's dilemma game where players are allowed to establish new interactions with others. By employing a simple coevolutionary rule entailing only two crucial parameters, we find that different selection criteria for the new interaction partners as well as their number vitally affect the outcome of the game. The resolution of the social dilemma is most probable if the selection favors more successful players and if their maximally attainable number is restricted. While the preferential selection of the best players promotes cooperation ir…
We study a spatial, one-shot prisoner's dilemma (PD) model in which selection operates on both an organism's behavioral strategy (cooperate or defect) and its choice of when to implement that strategy across a set of discrete time slots. Cooperators evolve to fixation regularly in the model when we add time slots to lattices and small-world networks, and their portion of the population grows, albeit slowly, when organisms interact in a scale-free network. This selection for cooperators occurs across a wide variety of time slots and it does so even when a crucial condition for the evolution of …
In this work we study a weak Prisonerâs Dilemma game in which both strategies and update rules are subjected to evolutionary pressure. Interactions among agents are specified by complex topologies, and we consider both homogeneous and heterogeneous situations. We consider deterministic and stochastic update rules for the strategies, which in turn may consider single links or full context when selecting agents to copy from. Our results indicate that the co-evolutionary process preserves heterogeneous networks as a suitable framework for the emergence of cooperation. Furthermore, on those networ…
Punishment and partner switching are two well-studied mechanisms that support the evolution of cooperation. Observation of human behaviour suggests that the extent to which punishment is adopted depends on the usage of alternative mechanisms, including partner switching. In this study, we investigate the combined effect of punishment and partner switching in evolutionary prisoner's dilemma games conducted on a network. In the model, agents are located on the network and participate in the prisoner's dilemma games with punishment. In addition, they can opportunistically switch interaction partn…
Cooperation in one-shot anonymous interactions is a widely documented aspect of human behaviour. Here we shed light on the motivations behind this behaviour by experimentally exploring cooperation in a one-shot continuous-strategy Prisoner's Dilemma (i.e. one-shot two-player Public Goods Game). We examine the distribution of cooperation amounts, and how that distribution varies based on the benefit-to-cost ratio of cooperation (b/c). Interestingly, we find a trimodal distribution at all b/c values investigated. Increasing b/c decreases the fraction of participants engaging in zero cooperation …
Cooperation is ubiquitous in nature, but explaining its existence remains a central interdisciplinary challenge. Cooperation is most difficult to explain in the Prisoner's Dilemma game, where cooperators always lose in direct competition with defectors despite increasing mean fitness. Here we demonstrate how spatial population expansion, a widespread natural phenomenon, promotes the evolution of cooperation. We engineer an experimental Prisoner's Dilemma game in the budding yeast Saccharomyces cerevisiae to show that, despite losing to defectors in nonexpanding conditions, cooperators increase…
In human societies the probability of strategy adoption from a given person may be affected by the personal features. Now we investigate how an artificially imposed restricted ability to reproduce, overruling ones fitness, affects an evolutionary process. For this purpose we employ the evolutionary prisoner's dilemma game on different complex graphs. Reproduction restrictions can have a facilitative effect on the evolution of cooperation that sets in irrespective of particularities of the interaction network. Indeed, an appropriate fraction of less fertile individuals may lead to full supremac…
Evolutionary prisoner's dilemma games are studied with players located on square lattice and random regular graphs defining four neighbors for each one. The players follow one of the three strategies: tit-for-tat, unconditional cooperation, and defection. The simplified payoff matrix is characterized by two parameters: the temptation $b$ to choose defection, and the cost $c$ of inspection reducing the income of tit-for-tat. The strategy imitation from one of the neighbors is controlled by pairwise comparison at a fixed level of noise. Using Monte Carlo simulations and the extended versions of …
Recent work has revealed a new class of "zero-determinant" (ZD) strategies for iterated, two-player games. ZD strategies allow a player to unilaterally enforce a linear relationship between her score and her opponent's score, and thus achieve an unusual degree of control over both players' long-term payoffs. Although originally conceived in the context of classical, two-player game theory, ZD strategies also have consequences in evolving populations of players. Here we explore the evolutionary prospects for ZD strategies in the Iterated Prisoner's Dilemma (IPD). Several recent studies have foc…
Here we study the effects of adopting different strategies against different opponent instead of adopting the same strategy against all of them in the prisoner dilemma structured in well-mixed populations. We consider an evolutionary process in which strategies that provide reproductive success are imitated and players replace one of their worst interactions by the new one. We set individuals in a well-mixed population so that network reciprocity effect is excluded and we analyze both synchronous and asynchronous updates. As a consequence of the replacement rule, we show that mutual cooperatio…
The prisoner's dilemma game is the most known contribution of game theory into social sciences. Here we describe new implications of this game for transactional and transformative leadership. While the autocratic (Stackelberg's) leadership is inefficient for this game, we discuss a Pareto-optimal scenario, where the leader L commits to react probabilistically to pure strategies of the follower F, which is free to make the first move. Offering F to resolve the dilemma, L is able to get a larger average pay-off. The exploitation can be stabilized via repeated interaction of L and F, and turns to…
Recently, the eco-evolutionary game theory which describes the coupled dynamics of strategies and environment have attracted great attention. At the same time, most of the current work is focused on the classic two-player two-strategy game. In this work, we study multi-strategy eco-evolutionary game theory which is an extension of the framework. For simplicity, we'll focus on the voluntary participation Prisoner's dilemma game. For the general class of payoff-dependent feedback dynamics, we show the conditions for the existence and stability of internal equilibrium by using the replicator dyna…
The theory of repeated games analyzes the long-term relationship of interacting players and mathematically reveals the condition of how cooperation is achieved, which is not achieved in a one-shot game. In the repeated prisoner's dilemma (RPD) game with no errors, zero-determinant (ZD) strategies allow a player to unilaterally set a linear relationship between the player's own payoff and the opponent's payoff regardless of the strategy that the opponent implements. In contrast, unconditional strategies such as ALLD and ALLC also unilaterally set a linear payoff relationship. Errors often happe…
Holding on to one's strategy is natural and common if the later warrants success and satisfaction. This goes against widespread simulation practices of evolutionary games, where players frequently consider changing their strategy even though their payoffs may be marginally different than those of the other players. Inspired by this observation, we introduce an aspiration-based win-stay-lose-learn strategy updating rule into the spatial prisoner's dilemma game. The rule is simple and intuitive, foreseeing strategy changes only by dissatisfied players, who then attempt to adopt the strategy of o…
Aging is always present, tailoring our interactions with others and postulating a finite lifespan during which we are able to exercise them. We consider the prisoner's dilemma game on a square lattice, and examine how quenched age distributions and different aging protocols influence the evolution of cooperation when taking the life experience and knowledge accumulation into account as time passes. In agreement with previous studies, we find that a quenched assignment of age to players, introducing heterogeneity to the game, substantially promotes cooperative behavior. Introduction of aging an…
We study the evolution of cooperation in the prisoner's dilemma game, whereby a coevolutionary rule is introduced that molds the random topology of the interaction network in two ways. First, existing links are deleted whenever a player adopts a new strategy or its degree exceeds a threshold value, and second, new links are added randomly after a given number of game iterations. These coevolutionary processes correspond to the generic formation of new and deletion of existing links that, especially in human societies, appear frequently as a consequence of ongoing socialization, change of lifes…
We study the problem of the emergence of cooperation in the spatial Prisoner's Dilemma. The pioneering work by Nowak and May showed that large initial populations of cooperators can survive and sustain cooperation in a square lattice with imitate-the-best evolutionary dynamics. We revisit this problem in a cost-benefit formulation suitable for a number of biological applications. We show that if a fixed-amount reward is established for cooperators to share, a single cooperator can invade a population of defectors and form structures that are resilient to re-invasion even if the reward mechanis…
When the interactions between cooperators (C) and defectors (D) can be partially avoided within a population, there may be an overall enhancement of cooperation. One example of such screening mechanism occurs in the presence of risk-averse agents (loners, L) that are neutral towards others, i.e., both L and its opponent, whatever its strategy, receive the same payoff. Their presence in the Prisoner's Dilemma (PD) game sustains the coexistence of cooperators and defectors far beyond the level attained in their absence. Another screening mechanism is a heterogeneous landscape obtained, for examp…
In spatial games players typically alter their strategy by imitating the most successful or one randomly selected neighbor. Since a single neighbor is taken as reference, the information stemming from other neighbors is neglected, which begets the consideration of alternative, possibly more realistic approaches. Here we show that strategy changes inspired not only by the performance of individual neighbors but rather by entire neighborhoods introduce a qualitatively different evolutionary dynamics that is able to support the stable existence of very small cooperative clusters. This leads to ph…
We investigate evolutionary dynamics of altruism with long-range interaction on a cycle. The interaction between individuals is described by a simplified version of the prisoner's dilemma (PD) game in which the payoffs are parameterized by $c$, the cost of a cooperative action. In our model, the probabilities of the game interaction and competition decay algebraically with $r_{AB}$, the distance between two players $A$ and $B$, but with different exponents: That is, the probability to play the PD game is proportional to $r_{AB}^{-α}$. If player $A$ is chosen for death, on the other hand, the p…
We will study a population of individuals playing the infinitely repeated Prisoner's Dilemma under replicator dynamics. The population consists of three kinds of individuals using the following reactive strategies: ALLD (individuals which always defect), ATFT (almost tit-for-tat: individuals which almost always repeat the opponent's last move) and G (generous individuals, which always cooperate when the opponent cooperated in the last move and have a positive probability $q$ of cooperating when they are defected). Our aim is studying in a mathematically rigorous fashion the dynamics of a simpl…
