AI 中文总结
该研究开发结合代谢缩放原理的空间显式森林群落动力学框架，推导树大小分布解析解，揭示其稳态幂律 regime 特征，建立大小分布缩放指数与森林状况的关联，为森林缩放规律提供统一动力学视角。
AI 中文摘要
关联物种大小、丰度和资源可获得性的缩放关系是生态学中最稳健的经验规律之一。然而，这些群落水平规律如何从生态过程中产生的机理解释仍难以捉摸。在此，我们通过开发一个最小化的空间显式森林群落动力学框架来解决这一缺口，该框架结合了种子扩散、受局地光可获得性限制的生长、局地竞争以及基于代谢缩放原理的全局资源约束。通过推导树大小分布的解析解，我们表明其稳态呈现出两种不同的幂律 regime，其指数由资源和空间竞争的相对强度控制。这些 regime 之间的交叉由种子注入与局地资源可获得性之间的相互作用决定，建立了大小分布的缩放指数与森林状况之间的明确关联。最后，我们表明边界干扰可打破竞争物种之间的生态平衡，并诱导传播至森林深处的效应，远超出单株植物的扩散范围。总之，这些结果为森林缩放规律提供了统一的动力学视角，具有广泛生物群落的潜在应用价值。
英文摘要
Scaling relations linking species size, abundance, and resource availability are among the most robust empirical regularities in ecology. However, a mechanistic explanation for how these community-level laws emerge from ecological processes remains elusive. Here, we address this gap by developing a minimal spatially explicit dynamical framework for forest communities that incorporates seed dispersal, growth limited by local light availability, local competition, and global resource constraints grounded in metabolic scaling principles. By deriving an analytical solution for the tree-size distribution, we show that its stationary state exhibits two distinct power-law regimes whose exponents are controlled by the relative strength of resource and spatial competition. The crossover between these regimes is set by the interplay between seed injection and local resource availability, establishing an explicit link between the scaling exponent of the size distribution and forest condition. Finally, we show that boundary disturbances can break the ecological balance between competing species and induce effects that propagate deeply into the forest bulk, far beyond the single-plant dispersal range. Together, these results provide a unifying dynamical perspective on forest scaling laws with potential applications to a broad range of biological communities.
Comments14 pages, 4 figures
